Gramma, a Galapagos tortoise that was the oldest living animal at the San Diego Zoo, has died at an estimated 141 years of ...

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Beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died at about 141, the zoo says, adding that ...

Gramma, the beloved Galápagos tortoise and the San Diego Zoo’s oldest resident, has died at an estimated age of 141. Known ...

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Gramma, a Galapagos tortoise that was the oldest living animal at the San Diego Zoo, has died at an estimated 141 years of ...

Gramma arrived at the San Diego Zoo sometime between 1928 and 1931, and was supposed to be several decades old at the time.