But now Gramma, a Galápagos tortoise, has eaten her last cactus fruit, the San Diego Zoo reported. She arrived in San Diego ...

Gramma, the beloved Galápagos tortoise and the San Diego Zoo’s oldest resident, has died at an estimated age of 141. Known ...

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Gramma, a Galápagos tortoise and the oldest resident of the San Diego Zoo, died Nov. 20. Her estimated 141-year lifespan spanned two world wars and 20 U.S. presidents.

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